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Impacts of low-tech restoration methods on soil, vegetation, and livestock grazing in Montana's sagebrush steppe
(Montana State University - Bozeman, College of Agriculture, 2024) Lowing, Nicholas Allen; Chairperson, Graduate Committee: Bok Sowell
In the semi-arid rangelands of western North America, water is a limiting factor for plant growth. In Montana's high elevation rangelands, snowmelt is the primary source of water for ephemeral streams and mesic meadows. Wildlife and livestock rely on these areas especially when surrounding uplands have dried. Less snowpack and early melt times associated with climate change pose a threat to forage resources in this region. In 2018 and 2019, small rock restoration structures were constructed in seven drainages in southwest Montana with the intention of increasing resiliency in these systems. In this study we compare stream reaches with restoration structures to reaches without structures to assess the impacts of this technique on soil moisture, plant cover, diversity, evenness, and production. We observed cattle grazing within and adjacent to treated drainages to determine how they use the landscape. We used soil moisture probes to measure soil water content once per month in June-September of 2021 and 2022. We used 0.5m2 frames to estimate plant cover, diversity, and evenness June-September in both years. At the end of each growing season, vegetation frames were clipped and weighed to estimate production. Soil and vegetation sampling occurred in 54 reaches across 7 drainages. We observed cattle near four treated drainages in mornings and evenings to assess whether cattle prefer mesic or upland areas. Cattle observations occurred in July and August of 2021 and 2022. Using linear fixed effects models, we detected no differences (p > or = 0.05) in soil moisture between treated and untreated reaches during any sampling period. We detected no differences in vegetation cover, richness, diversity, or evenness in 92% of sampling periods. No differences in plant production were detected in 2021 or 2022. Cattle used mesic areas in greater proportion to their abundance on the landscape in both years (p < or = 0.0001). Our results indicate that these structures have not yet had major impacts on soil moisture or vegetation metrics. However, our results indicate that mesic meadows are an important resource for grazing cattle in southwest Montana's high- elevation rangelands, supporting the idea that these areas warrant restoration efforts.
Supplement intake behavior of heifers grazing late summer dryland pasture
(Montana State University - Bozeman, College of Agriculture, 2022) McClain, Tyrell Phillip; Chairperson, Graduate Committee: Timothy DelCurto; This is a manuscript style paper that includes co-authored chapters.
Two studies were conducted to evaluate supplement intake and intake behavior of Rumax BoviBox protein supplements. For both studies, individual supplement intake, time spent at the feeder, and frequency of visits was measured using a SmartFeed Pro self-feeder system. In study 1, we examined the effects of free choice loose mineral salt on protein block supplement intake behavior of first-calf heifers. Heifers were stratified by weight, and, within stratum, randomly allotted to one of two supplementation treatments: 1) free-choice access to protein block supplement (30% CP) with access to loose mineral and salt; and 2) free-choice access to protein block supplement (30% CP) with no access to loose mineral and salt for a 42-day performance study. Individual animal was considered the experimental unit. No differences were observed for total supplement intake or supplement intake expressed as grams per kg body weight (BW; P > 0.05). In summary, availability of loose mineral salt did not have an impact on protein block supplement intake. In study 2, we evaluated Rumax BoviBox versus Rumax BoviBox HM protein supplements on supplement intake, intake behavior, body weight (BW), and body condition score (BCS) change of yearling heifers grazing dryland pastures during late summer. Heifers (428 kg) were stratified by BW and within stratum randomly assigned to one of two supplementation treatments: 1) free-choice access to Rumax BoviBox protein block supplement (30% CP, 23% salt; n = 29); and 2) free-choice access to Rumax BoviBox HM high magnesium protein block supplement (28.7% CP, 23% salt; n = 30). Supplement intake rate (g x min -1) displayed a treatment effect (P < 0.01) indicating that heifers in the Rumax BoviBox treatment had lower intake rate compared to Rumax BoviBox HM supplemented heifers. In conclusion, there were only minor differences in intake behavior and animal performance observed in both trials.
The influence of heifer post-weaning residual feed intake on subsequent production, reproduction, grazing behavior, supplement intake behavior, dry matter intake and milk production of Black Angus beef cattle
(Montana State University - Bozeman, College of Agriculture, 2021) Parsons, Cory Todd; Chairperson, Graduate Committee: Timothy DelCurto; Julia Dafoe, Samuel Wyffels, Tim DelCurto, Darrin Boss and Megan Van Emon were co-authors of the article, 'Impacts of heifer post-weaning residual feed intake classification on reproductive and performance measurements of first, second and third parity Angus beef' in the journal 'Translational animal science' which is contained within this dissertation.; Julia Dafoe was a co-author of the article, 'The influence of residual feed intake and cow age on body weight and body condition change, supplement intake, resource use, and grazing behavior of beef cattle winter grazing mixed-grass rangelands' in the journal 'Animals' which is contained within this dissertation.; Julia Dafoe, Samuel Wyffels, Tim DelCurto and Darrin Boss were co-authors of the article, 'The influence of residual feed intake and cow age on dry matter intake post-weaning and peak lactation of Black Angus cows' in the journal 'Animals' which is contained within this dissertation.
Residual feed intake (RFI) is a recognized measure of biological efficiency in beef cattle. However, RFI determination is expensive, time consuming and not well studied in beef cattle fed forage-based diets, nor how post-weaning RFI influences grazing behavior, supplement intake behavior or dry matter intake at different ages and physiological stages of production. Thus, the objectives of this research were to investigate how post-weaning RFI influences reproductive and productive performance, grazing behavior, supplement intake behavior as well as dry matter intake at different ages and physiological stages of production. A commercial herd of black Angus cows were utilized to conduct three experimental studies. Results indicate that classification of RFI for replacement heifers had little to no effect on subsequent beef cattle production and reproductive efficiency through the weaning of the 3rd calf. Subtle differences were denoted for cow Julian birth dates based on RFI classification and conception of 1st calf-heifers categorized as low RFI. However, heifer post-weaning RFI had little effect on subsequent cow performance (BW or BCS), grazing behavior, supplement intake behavior, and resource use. In addition, cow age significantly influenced cow performance, grazing behavior, supplement intake behavior, and resource use. We also observed high individual variability in grazing site selection, suggesting that individual-level factors may be driving grazing resource use and grazing behavior. Heifer post-weaning RFI did not influence mature cow dry matter intake, and this was consistent for both lactating and non-lactating beef cows. In contrast, cow age did correspond to quadratic increases of DMI and intake rates of mature cows. However, when DMI was expressed as g x kg body weight-1 no differences were observed with respect to cow age in lactating and non-lactating cows. Milk production was influenced by heifer post-weaning RFI for 2/3 and 5/6-yr old cows. Therefore, our studies suggests that selection for low RFI females would not impact overall herd productivity of cattle on foraged-based production systems, and that cow age has more of an impact on resource use, supplement intake, and grazing behavior, as well as DMI than heifer post-weaning RFI while grazing dormant-season mixed-grass prairie rangelands.
Effects of grazing after wildfire on soil health in eastern Montana
(Montana State University - Bozeman, College of Agriculture, 2021) Hanson, Joshua Todd; Chairperson, Graduate Committee: Clayton B. Marlow; Clayton B. Marlow, Kurt O. Reinhart, Lance T. Vermeire and Sam A. Wyffels were co-authors of the article, 'Effects of grazing after wildfire on indicators of soil health in eastern Montana' submitted to the journal 'Fire ecology' which is contained within this thesis.
Rangelands are resilient to grazing and fire. However, the resilience of rangelands may be degraded by livestock grazing too soon after wildfire. Due to the growing interest in soil health and its link to sustainable grazing, following a large wildfire (109,346 ha, Lodgepole complex in 2017) we tested the effect of grazing (grazed verses no grazing) on three indicators of soil health in ponderosa pine savvanas of the northern Great Plains. We measured indicators of soil hydrologic function (i.e., soil hydraulic conductivity), properties related to nutrient cycling (i.e., soil organic matter, plant available nutrients, pH), and soil structure (i.e., aggregate stability) in 2019 and 2020. Grazing occurred two out of three years following the fire. Most indicators of soil health were not appreciably affected by grazing post-fire. However, we detected a marginally significant negative effect of grazing on soil organic matter. Specifically, soil organic matter was 7% greater in ungrazed than grazed areas. No other grazing treatment effects were detected. Several soil health metrics varied between sampling years. Our results suggest that grazing(two out of the three years) following fire is unlikely to negatively affect many indicators of soil health of ponderosa pine savannas in the northern Great Plains. The increased organic matter observed by resting did not have an appreciable impact on the other soil health metrics. These findings suggest that soil health indicators are relatively resilient to grazing after wildfire and have implications for grazing policy post-fire.
Evaluating grazing and defoliation effects on ponderosa pine grassland following the lodgepole complex wildfire
(Montana State University - Bozeman, College of Agriculture, 2020) Williams, Amanda Rae; Chairperson, Graduate Committee: Clayton B. Marlow; Lance T. Vermeire, Richard C. Waterman and Clayton B. Marlow were co-authors of the article, 'Evaluating grazing effects on ponderosa pine grassland following the lodgepole complex wildfire' submitted to the journal 'Rangeland ecology and management' which is contained within this thesis.; Lance T. Vermeire, Richard C. Waterman, Clayton B. Marlow were co-authors of the article, 'Season of defoliation effects on ponderosa pine grassland following the lodgepole complex wildfire' submitted to the journal 'Rangeland ecology and management' which is contained within this thesis.
Previous research indicated rangelands need rest from grazing after fire, while others show grazing the first year following fire has no negative effects on the plant community. This caused uncertainty around post-fire grazing management. In July 2017, the Lodgepole Complex fire burned ponderosa pine grasslands of the northern Great Plains, including areas burned in 2003 for fuels mitigation. We examined effects of post-fire grazing and season of defoliation. For the grazing study we wanted to determine 1) plant community response to grazing or rest the first growing season after fire in ponderosa pine grassland communities, and 2) whether prescribed fire alters plant community response to subsequent wildfire. For the season of defoliation study we wanted to determine 1) timing of defoliation effects on the plant community one growing season after fire and 2) whether defoliation effects are altered by prescribed fire preceding the fire. Eight exclosures (25 x 15 m) were built, 4 reburned sites and 4 wildfire sites. A non-grazed (15 x 10 m) section inside each exclosure was paired with a grazed section outside the exclosure. Plots (5 x 10 m) were mowed in the exclosure to 10 cm in June, July, or August, or not mowed during 2018. In 2019, biomass samples were clipped at peak production, with species composition and diversity measured by point-intercept transects. We observed a trend for reduced cool-season grasses (P =0.0675) and annual grasses (P =0.0071) if defoliated earlier; a trend for reduced forbs (P =0.0699) if defoliated later; and reduced total current-year biomass (P =0.0362) if defoliated. Functional group composition was not changed, but some individual species were shifted due to fire history. The grazing study only showed a trend for greater old dead biomass on non-grazed sites (P = 0.0600), higher composition of forbs on reburn sites (P = 0.0324), and a trend for a higher composition of Psoralidium tenuiflorum (Pursh) Rydb. on reburn sites (P = 0.0559). Results indicate mowing the first year following fire shifts the plant community, but the community is resistant to post-fire grazing. Prescribed fire 14 years before wildfire had small impacts on community composition.
Native pollinators: the effects of livestock grazing on Montana rangelands
(Montana State University - Bozeman, College of Agriculture, 2019) Blanchette, Gabrielle Elizabeth; Chairperson, Graduate Committee: Craig Carr; Michael A. Ivie (co-chair)
Although native pollinators on rangelands serve, in part, as food at higher trophic levels, their primary ecosystem function is pollination. With 70% of western U.S. rangelands grazed by livestock, understanding how grazing affects native pollinators is a key component to managing rangelands, yet it is not well understood. In this study, I investigated how cattle grazing influences both diversity and abundance of native pollinators, including bees and wasps, syrphid flies, butterflies, and moths at two research locations for six to 10 weeks during the spring of 2016, 2017, and 2018. The first site, near Sidney, MT, served to understand if pollinators were more closely associated with active cattle grazing or rested pastures. The second site, near Roundup, MT, served to understand if pollinators were more associated with pastures either enrolled or not enrolled in the Sage Grouse Initiative, or pastures that had not experienced livestock grazing in previous seven years. Colored pan traps were deployed weekly in each treatment at each site. In addition to pollinator collections, weekly measurements of vegetation via Daubenmire frame were also collected. I collected 17,078 specimens at Sidney and 13,683 specimens at Roundup. My results suggest that in drier sagebrush landscapes, native pollinators are positively to neutrally associated with pastures moderately grazed by livestock. However, in mixed grass prairie landscapes that receive high precipitation, rest-rotational grazing does not appear to have a significant effect on primary native pollinators.
Dormant season grazing of northern mixed grass prairies: effect of supplementation and winter environmental conditions on beef cattle grazing behavior, residual vegetation conditions and variation in supplement intake
(Montana State University - Bozeman, College of Agriculture, 2019) Wyffels, Samuel Aaron; Chairperson, Graduate Committee: Lance McNew; Janice G. P. Bowman (co-chair); Lance B. McNew, Janice G.P. Bowman, Mark K. Petersen, Darrin L. Boss and Bok F. Sowell were co-authors of the article, 'Dormant season grazing of northern mixed grass prairies: the effect of supplementation strategies on heifer resource utilization and vegetation use' submitted to the journal 'Rangeland ecology & management' which is contained within this thesis.; Lance B. McNew, Janice G.P. Bowman, Mark K. Petersen, Darrin L. Boss and Bok F. Sowel were co-authors of the article, 'Dormant season grazing on northern mixed grass prairie agroecosystems: does protein supplementation and individual animal variation impact beef cattle resource use, vegetation and residual cover for wildlife' submitted to the journal 'Rangeland ecology & management' which is contained within this thesis.; Lance B. McNew, Janice G.P. Bowman, Mark K. Petersen, Darrin L. Boss and Bok F. Sowell were co-authors of the article, 'The influence of age and environmental conditions on supplement intake by beef cattle winter grazing a northern mixed-grass rangeland in Montana' submitted to the journal 'Rangeland ecology & management' which is contained within this thesis.
Dormant season grazing reduces reliance on harvested feeds, but typically requires protein supplementation to maintain animal performance and vegetation utilization across the landscape. However, information relating supplementation strategies and supplement intake behavior to individual grazing behavior and resource utilization on dormant forage is lacking. Thus, the intent of this research is to examine cattle resource utilization, supplement intake behavior, residual cover of vegetation and utilization on rangelands grazed during the dormant season. One hundred weaned heifer calves were randomly selected and placed into one of two supplementation treatments in each of 2 years (50 heifers/treatment/year); one receiving a free access 62% crude protein self-fed mineral/protein concentrate, and the other receiving a daily hand-fed 20% crude protein cake while grazing December through March. Additionally, a commercial herd of 300 bred cows ranging in age from 1- to 12-yr-old were provided a 30% crude protein self-fed supplement with 25% salt to limit intake in a SmartFeed Pro self-feeder system to measure individual animal supplement intake from November to January in each of 2 years. In both grazing trials, transects were randomly located within each pasture for measuring vegetation composition, production and quality, canopy cover and visual obstruction readings pre and post grazing. Grazing locations were monitored for individuals with Global Positioning System collars containing head position sensors that record daily space use and location of grazing activities. Data sets were used to quantify space use with generalized linear models to assess cattle resource utilization and supplementation behavior. Cattle provided the hand-fed cake selected grazing location near supplement delivery sites and spent less time grazing per day than self-fed supplemented cattle. Substantial amounts of herd-level variability in both studies suggests individual attributes are major drivers in cattle resource use. Supplement treatment and grazing intensity had little impact on residual vegetation conditions, however, the timing of grazing and year did affect the response of residual vegetation to grazing. Younger cattle consumed more supplement with less variability than older aged cattle. This research provides multidimensional insight to stakeholders concerning grazing behavior and the ecological impacts of late season use on Montana rangelands.
Livestock depredation by grizzly bears on Forest Service grazing allotments in the Greater Yellowstone Ecosystem
(Montana State University - Bozeman, College of Agriculture, 2017) Wells, Smith Laura; Chairperson, Graduate Committee: Lance McNew
Grizzly bear population growth and range expansion over the last several decades in the Greater Yellowstone Ecosystem (GYE) has led to increased human-bear conflicts, including livestock depredation on public land grazing allotments. A better understanding of patterns and relationships between grazing allotment management and grizzly bear depredation of livestock is needed for adaptive, sustainable management in the ecosystem. Historic U.S. Forest Service and National Park Service livestock grazing records, grizzly bear habitat attributes, and documented livestock depredations by grizzly bears were collated for 316 public land grazing allotments within the grizzly bear Demographic Monitoring Area (DMA) during 1992-2014. Spatio-temporal relationships between annual livestock depredation counts and grazing allotment characteristics were modeled for each allotment during the study period at two spatial extents, representing daily and annual grizzly bear activity areas. As the Yellowstone grizzly population expanded during the last several decades, more public land grazing allotments were exposed to potential livestock-grizzly bear interactions and results indicated that both livestock stocking and grizzly bear habitat characteristics in and around allotments were related to documented depredations during 1992-2014. Annual numbers of livestock and grizzly bear density on allotments had a large, positive effect on average livestock depredation event counts. Allotment size and summer grazing both were related to higher depredation event counts while the presence of bulls and/or horses was related to lower counts. Allotments with less rugged terrain, lower road density, relatively higher vegetative primary productivity, greater amounts of whitebark pine, and further from forest edge on average were associated with higher average livestock depredation event counts. Managers and livestock producers could use these results to support adaptive management approaches and long-term planning such as increasing herd supervision, especially in areas with quality grizzly bear habitat and high grizzly bear density, or altering grazing management strategies and grazing locations to limit potential livestock depredation events. Results provide insight into historic livestock-grizzly bear conflicts on public lands in a large, complex ecosystem and although challenging, results could support cooperative management strategies to sustain the grizzly bear population and livestock operations in the GYE.
Grazing, feed intake, and milk production differences in beef cows of varying efficiency classification and size
(Montana State University - Bozeman, College of Agriculture, 2018) Williams, Alyson Rose; Chairperson, Graduate Committee: Timothy DelCurto; C.T. Parsons, J.G.P. Bowman, D.L. Boss and T. DelCurto were co-authors of the article, 'The influence of weaning weight ratio and cow size on feed intake behavior, milk yield, and milk constituents' submitted to the journal 'Journal of animal science' which is contained within this thesis.; S.A. Wyffels, C.T. Parsons, J.G.P. Bowman, D.L. Boss and T. DelCurto were co-authors of the article, 'The influence of weaning weight ratio and cow size on winter grazing and supplement intake behavior' submitted to the journal 'Journal of animal science' which is contained within this thesis.
Metrics that identify beef cow efficiency in extensive rangeland environments has long been desired in both beef cattle research and production settings. However, research is limited relative to matching cattle metrics to western production systems. Previous research has suggested the ratio of calf weight weaned to cow weight, or weaning weight ratio, is an accurate estimate of cow efficiency. Furthermore, milk production has been attributed to influence calf pre-weaning average daily gain. Few studies have evaluated how cow type or cow characteristics influence grazing distribution and supplement consumption on native landscapes. The objectives of this study were to assess cow feed intake, grazing behavior, supplement intake, and milk production differences between cows of differing weaning weight ratio and body weight classification. Individual cow average daily feed consumption, average daily feeding bout duration, number of visits per day, and total time spent eating per day were collected during a feed intake period and a supplement intake period. Total distance traveled and time spent grazing were collected on individual cows with global positioning system collars. Milk samples were analyzed for fat, solids not fat, total solids, protein, and lactose content, and milk yield was calculated as the change in calf weight following a weigh-suckle-weigh procedure. High weaning weight ratio cows consumed more feed when expressed on a g/kg bodyweight basis (P < 0.001) but did not differ in supplement consumption (P > 0.10). High weaning weight ratio cows produced more milk both in total kg (P < 0.03) and when expressed as /kg bodyweight (P < 0.001). Heavy bodyweight cows tended to consume more feed than light bodyweight cows (P < 0.09) but neither bodyweight group ate more supplement than the other (P > 0.10). High weaning weight ratio cows spent an hour longer grazing per day than low weaning weight ratio cows (P < 0.02). Total distanced traveled per day did not differ (P > 0.10). Results suggest weaning weight ratio and cow size are not accurate metrics of cow efficiency and the use of weaning weight ratio may unintentionally favor cows with higher milk production and higher feed requirements.
Effects of livestock grazing management on the ecology of grassland birds and their predators in a northern mixed-grass prairie ecosystem
(Montana State University - Bozeman, College of Agriculture, 2018) Vold, Skyler Thomas; Chairperson, Graduate Committee: Lance McNew; Lance B. McNew and Lorelle I. Berkeley were co-authors of the article, 'Effects of livestock grazing management on the ecology of grassland birds in a northern mixed-grass prairie ecosystem' submitted to the journal 'Rangeland ecology & management' which is contained within this thesis.; Lance B. McNew and Lorelle I. Berkeley were co-authors of the article, 'Effects of livestock grazing management on the occupancy of mesocarnivores in a northern mixed-grass prairie ecosystem' submitted to the journal 'The journal of wildlife management' which is contained within this thesis.
Grassland birds have recently undergone substantial population declines throughout much of their historic ranges in North America. The majority of remaining grassland bird habitat is rangeland managed for livestock production, so grazing management has implications for grassland bird conservation efforts. Populations of mesocarnivores have recently seen range expansions and increased abundance as a result of anthropogenic extirpation of apex predators in grassland ecosystems. Mesocarnivores are often major predators of grassland birds and their nests, so considering the effects of management actions on mesocarnivore occupancy is important within grassland bird conservation efforts. I evaluated the relative effects of three livestock grazing systems on grassland bird abundance and mesocarnivore occupancy in a northern mixed-grass prairie ecosystem of eastern Montana, USA. During 2016-17, I conducted replicated point-count surveys at 150 locations on a 3,000-ha ranch managed with rest-rotation cattle grazing, and 155 locations on adjacent reference properties (4,300-ha) employing season-long or 2-pasture summer-rotational grazing systems to identify grazing management influences on grassland bird abundances. During 2016-17, I deployed remote cameras at 45 locations within rest-rotation grazing systems, and at 45 locations on reference properties to identity grazing management influences on mesocarnivore occupancy. Overall, there was no noticeable benefit of rest-rotation grazing on abundance or species diversity of grassland birds relative to season-long and summer-rotation grazing systems. Species-specific responses to livestock grazing system were found for each of three obligate grassland birds. Support for interactions between grazing system and local rangeland production potential were found, limiting the ability to recommend general livestock management practices for the benefit of grassland bird populations. Additionally, abundance of grasshopper sparrow (Ammodramus savannarum; beta = -0.10 + or = 0.03) and western meadowlark (Sturnella neglecta; beta = -0.09 + or = 0.03) showed a negative relationship with increasing stocking rate. Occupancy of mesocarnivores was highest in rest-rotation grazing systems, followed by season-long and summer-rotation systems, respectively, and showed a positive relationship with increasing stocking rate (beta = 1.64 + or = 1.10). Regional guidelines for livestock grazing management may be inappropriate in terms of grassland bird conservation efforts in the northern mixed-grass prairie.

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