College of Letters & Science

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The College of Letters and Science, the largest center for learning, teaching and research at Montana State University, offers students an excellent liberal arts and sciences education in nearly 50 majors, 25 minors and over 25 graduate degrees within the four areas of the humanities, natural sciences, mathematics and social sciences.

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Vegetation Distribution and Production in Rocky Mountain Climates—with Emphasis on Whitebark Pine
(International Workshop on Subalpine Stone Pines and Their Environment: the Status of Our Knowledge, 1992) Weaver, T.
The distribution and production of vegetation on the altitudinal J gradient (grassland-forest-alpine) was plotted against climatic parameters to evaluate hypothetical controlling factors. (1) Whitebark pine (Pinus albicaulis) is likely excluded from higher zones by a cool growing season or wind-induced drought. It is probably not excluded by low temperatures occurring during its hardening, hard, or dehardening seasons. (2) While the lower physiological limit of whitebark pine is probably set by drought its lower realized limit is directly set by subalpine fir (Abies lasiocarpa) and lodgepole pine (Pinus contorta) competitors and indirectly set by factors that control their distribution. (3) The upper limits for most other dominant species are probably set by growing season temperature. The lower limits are likely set by competition down to the cedar-hemlock (Thuja plicata/Tsuga heterophylla) zone and by drought in drier areas. (4) Production is strongly correlated (r 2 = 0.86) with growing season length (soil thawed season minus dry soil days). Multiplying season length by average temperature did not improve the growing season predictor, perhaps because vegetation at each altitude is especially adapted to temperatures in its zone.
Climates Where Stone Pines Grow, A Comparison
(International Workshop on Subalpine Stone Pines and Their Environment: the Status of Our Knowledge, 1992) Weaver, T.
------ Abstract-While stone pine climates are similar adapted to relatively moderate climates may be excluded from, species the ranges of congeners by more severe climates, and species with longer warm-moiBt growing seasons are probably more productive than congeners. Absolute low/summer average/absolute high temperatures for stone pines listed in order of increasing absolute low temperature are Pinus sibirica (-65/13/37 °C), P. pumila (-52.19/36 °C), P. koraiensis (-42/1s1/36 °C), P. albicaulis (-3419/29 °C), and P. cembra (-23/8/27 °C). The Walter drought index shows little stress in stone pine forests despite large differences in summer/winter precipitation: in order of increasing summer rainfall, precipitation is P. albicaulis (102.I 829mm), P. pumila (1421.264mm), P. sibirica (1871245=), P. cembra (3231616mm), P. koraieT18is (3941242mm). Estimated thawed-soil growing season increases from P. albicaulis (4.5mo), throughsP. pumila (4.6mo), P. sibirica (5.5mo), and P. cembra (6.3mo) to P. koraiensis (7.8mo); growing seasons of the first three trees could be shortened by drought.
Distribution of Exotic Plants in the N. Rocky Mountains by Environmental Type and Disturbance Condition
(Montana State Univeristy, 1989-06) Weaver, T.; Gustafson, D.; Lichthardt, J.; Woods, B.
This report lists seventy-three exotic species found in a systematic sampling of major environmental zones of the Rocky Mountains between the Canadian border and central Wyoming. For each exotic it states the regional distribution, the environmental types (HTs) it occupies (% constancy), the disturbance conditions (DCs) it occupies (% constancy), and its dominance (in terms of% frequency and% cover) in each cell of the HT x DC matrix. Park managers need to develop policy with respect to legally noxious weeds, forage grasses (eg Phleum pratense, Poa pratensis, Bromus inermis, and Dactylis glomerata), and forage legumes (eg Melilotus and Trifolium spp).
PLANTS ESTABLISHING IN ROCKY MOUNTAIN ENVIRONMENTS-- a manual for choosing native species for revegetation
(Montana State University, 1995) Weaver, T.; Gustafson, D; Lichthardt, J
Species which have established naturally on a disturbed site in a given environment-- climate and disturbance level (defined below)-- are good candidates for revegetation plantings in that environment. On this basis we recommend native plants (grasses, forbs, and shrubs) for revegetation plantings, if they occur on at least half of the sites sampled in the environmental type and cover at least 1% of the ground there. We also list exotic plants establishing on once disturbed roadside sites; if these plants do not invade native vegetation they might, under some circumstances, be used for revegetation The environmental types considered include dry grasslands (BOGR/STCO and AGSP/BOGR), moist grasslands (FESC/FEID and FEID/AGCA) sagebrush (ARAR/FEID and ARTRVAS/FEID) , warm dry forests (PSME/ SYAL and PSME/PHMA), warm moist forests (POTR/CARU, THPL/OPHO, TSHE/CLUN, ABLA/CLUN), cool forests (ABLA/XETE, ABLA/ARCO, and ABLA/VACC), mountain meadows (FEID/AGCA, listed above) and alpine (DESC/CARX) . In each environment plant performance is contrasted across five disturbance types: continually disturbed types (roadshoulders and the adjacent ditch slope), once disturbed sites (roadcuts with organic matter removed and cleared right-of-way without organic matter removal), and undisturbed late seral sites.
Changes in soils along a vegetational (altitudinal) gradient of the northern Rocky Mountains
(Soil Science Society of America, 1979) Weaver, T.
As one moves from the warm dry plains of eastern Montana to the cool moist peaks of the northern Rocky Mountains he might pass through a series of native vegetation types: Bouteloua gracilis, Agropyron spicatum, Featuca idahoensis, and Festuca scabrella grasslands; Pinua ponderosa, Pseudotsuga menziesii, and Abies lasiocarpa forests; and alpine tundra (Kuchler 1964, Muggler and Handl 1974, Pfister et al. 1977). It is commonly observed that when one moves up a vegetational gradient he moves up a soils gradient (e.g. Eyre 1963, Whittaker et al. 1968, Hanawalt and Whittaker 1976 and 1977). In the northern Rocky Mountains, Thorp (1931, N Wyoming) observed that organic matter increased, that pH decreased, that the depth to free lime increased and that the thickness of A- and B-horizons increased as he moved up a vegetational gradient similar to that described above. The same trends, as well as a tendency for nutrients to become most available at the grassland-forest boundary, were observed along a similar vegetation gradient in British Columbia (Spilsbury and Tisdale 1944). Such trends correlate well with broad groups in the 1938 Soil Taxonomy (Agricultural Experiment Station 1964 and Nimlos 1963) as well as in the 1977 Soil Taxonomy (Weaver 1978). The objects of this paper are I) to describe the change in soils observed along this gradient in more detail, 2) to consider their genesis briefly, and 3) to consider their importance to plants.
Root distribution and soil water regimes in nine habitat types of the northern Rocky Mountains
(Colorado State University, 1977) Weaver, T.
Root distribution and the annual cycle of' soil water availability were measured in nine habitat types of the northern Rocky Mountains. Water stress periods became progressively longer under Abies lasiooarpa forests, Populus trerrruloides groves, Pseudotsuga menziesii forests, Festuoa idahoensis grasslands, Artemisia tridentata shrublands, and Agropyron spioatum grasslands. Water stress periods were longer under Pseudotsuga forests than under adjacent logged areas. Live feeder root biomass (1) was similar under grassland, shrubland, and forest types, (2) increased within a vegetation type with altitude, and (3) decreased at a site with depth. Seral grasslands had less live feeder root biomass than forests in the same habitat type, but climax grasslands and forests were similar in root biomass.
Cone production in Pinus albicaulis Forests
(Inland Mountain West Symposium, 1985-08) Weaver, T.; Forcella, F
Whitebark pine cone production was estimated for a 6 to 8 year period in each of 29 stands widespread in.the northern.Rocky Mountains. 1) One-time sampling was possible since the estimate was m2de by multiplying the number of branches perm by an estimate of annual cone production made from counts' of cone lets, mature cones, or cone scars on successively older annual increments of those branches. 2) Average cone production ranged from 0.3 to 3.6 cones·m^-2 ·yr^1 and from 22-270 seeds·m^-2·year^-1 . 3) Regression analysis was used to relate the variance observed to time and place. a) Year-to-year variation in the cone yield of branches, trees, and stands in a region appears to be both internally and externally controlled. Internal control is suggested by the fact that good cone years were usually preceded by poor cone years. While external control is indicated by significant correlations between growth and weather conditions, control is not dominated by the effect of any one factor or any particular developmental stage. b) Although cone production of the average branch varied significantly within 30 percent of the trees and within 48 percent of the stands observed, it did not vary significantly among stands. c) Regressions relating stand cone production to easily measured stand characteristics such as canopy cover, fallen cones, and/or stand size explain no more than 50 percent of the variance among stands.
Bibliography of Montana vegetation description
(1988) Burgeron, P.; Kratz, A.; Weaver, T.; Weidman, N.
Listed in alphabetical order by author are 549 references to literature that describes the native vegetation of Montana. This updates the 1965 list of Habeck and Hartley. A keyword subject index is included.
Biomass and productivity of the subalpine Pinus albicaulis — Vaccinium scoparium association in Montana, USA
(1977-10) Forcella, F.; Weaver, T.
Biomass of fifty conifer forests and nutrient exports associated with their harvest
(1977) Weaver, T.; Forcella, F.
Biomasses of climax Rocky Mountain forests studied ranged from less than 50 to more than 300 tons/ha. Total biomass was approximately 1.5 times the biomass of normally merchantable boles. When compared with conventional bole harvest, the nutrient exports associated with harvest of all aboveground parts in these stands would apparently be at least three times higher for nitrogen, six times higher for phosphorus, four times higher for potassium, and three times higher for calcium.

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